They are put on the pto shaft so that the inertia created by an implement like a bush hog's blades spinning doesn't keep making the transmission spin when you even push in the clutch. On older tractors it actually can power the transmission enough to keep you from stopping the tractor from moving for a few minutes, causing you to drive into a fence row or tree.
Most newer tractors have an indepentant pto and do not need an over running coupler. I don't know about a JD 4300. I would ask a dealer.